Bachelors: Reed College, Portland, OR, 1968, biology (one year at the University of York, England)
Diplom (masters): Technical University of Braunschweig, 1972, major: Microbiology, minors: Botany and Organic Chemistry
PhD: Technical University of Braunschweig, 1980, major: Microbiology, minors: Botany and Biochemistry; Thesis: "Relationships between morphology and metabolism in the dimorphic fungus Mycotypha (Mucorales)
Habilitation: Technical University of Braunschweig, 2003, Plant Pathology